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Controls

You can change any of these in Options → Controls → Key Binds → Transformer.

Key Action
R Transform between robot and vehicle mode
Z Get your guns out, or put them away
G Get your melee weapon out, or put it away
V Special ability (robot mode) or headlights (vehicle mode)

To shoot, get your guns out, keep your main hand empty and left-click. Hold it down to keep firing. For melee, get your weapon out and left-click to swing.

Shooting uses a little Energon, about one shot in four, and you can’t shoot once your Energon bar is down to two icons.

Character Controls
Optimus Prime, Megatron (with their saber) Hold left-click to charge, double-tap sprint to dash, right-click to block. See the Star Saber and Dark Star Saber
Dreadwing, Skyquake Hold right-click with your gun out to rev the cannon or minigun
Skyquake Hold left-click with the minigun out for continuous fire
Knockout, Vehicon Miner Hold left-click for a continuous saw (Knockout) or drill (Vehicon Miner) attack
Airachnid Crouch and press R in robot mode to switch to spider mode
Arcee, Vehicon Miner Crouch and press V in vehicle mode to toggle the hologram rider
Scraplet morph V burrows into the block you’re looking at. Left-click while hidden to peek out. Z switches heat vision on and off

Decepticons give their recruited Vehicons orders by crouching while looking at something: an enemy to attack it, a block to hold that spot or up to regroup. See Giving orders.